Mandrills live on a diet of fruit, seeds, bark, leaves, insects, palm nuts and small animals. They are called "seed predators" because they forage on seeds all day as they roam along the savannah. Mandrills are quadrupedal, arboreal and terrestrial. They are active during the day. Mandrills travel in multi-male multi-female groups in which the males have a strict hierarchy. Males possessively guard females that are in estrous. Head bobbing is common in this species. Males seven years and older scent mark which is a rare trait in old world monkeys.